Epaphus was the son of Zeus and Io and thus, Ceroessa's brother. With his wife, Memphis (or according to others, Cassiopeia), he had one daughter, Libya while some accounts added another one who bore the name Lysianassa. These daughters later became mothers of Poseidon's sons, Belus, Agenor and possibly, Lelex to the former and Busiris to the latter. In other versions of the myth, Epaphus was also called father of Thebe, mother of Aegyptusand Heracles by Zeus. Through these daughters, Epaphus was the ancestor of the "dark Libyans, and high-souled Aethiopians, and the Underground-folk and feeble Pygmies".